Edited sentences: role

At both study sites, M. senhousia played a double role: (1) as secondary substratum, enhancing the environmental structural complexity, and (2) as densely aggregated, living organisms, thus giving rise to a number of possible interactions with other invertebrates.

M. senhousia played a double role
M. senhousiaは2つの役割を演じた

enhancing the environmental structural complexity
環境の構造上の複雑さを高めること

densely aggregated
塊状に密集させた